Isatis cappadocica
Isatis cappadocica
Ssp alyssifolia: Has çivitotu; Ssp besseri: Çölemerik çivitotu; Ssp cappadocica: Peri çivitotu;
Ssp macrocarpa: Hakkari çivitotu; Ssp nurihakensis: Nurhak çiviti; Ssp steveniana: Oyali çivit;
Ssp subradiata: Meşe çivitotu
Perennial, often suffrutescent. Stems 20-70 cm. Leaves thin, like the stems usually pubescent or glabrous; basal leaves oblanceolate or obovate, entire or weakly dentate; median stem leaves oblong to linear-oblong or oblong-lanceolate, obtusely or acutely auricled. Flowers 4-5 mm. Fruits 10-30 x 5-20 mm, in Turkey obovate, elliptic, ovate, suborbicular, ovate-rhomboid or cuneate-oblong, shortly and densely hairy or glabrous, wing thin, usually much broader than the loculus, flat or undulate, loculus not transversely wrinkled, midrib slender.
An extremely polymorphic species whose limits and subdivision remain obscure. As circumscribed here, the species is equivalent to most of Boissier's inter-pretation of I. latisiliqua Stev. though it excludes I. latisiliqua itself, a distinctive species confined to Caucasia and distinguished from I. cappadocica by its hispid stem bases and transversely wrinkled loculus Desv.). The key below attempts to cope with most of the major variants found in Turkey; other taxa in the same complex occur in Caucasia and Iran . Examination of the type in Paris shows I. cappadocica to be the earliest name for the group.
1. Fruits 10-12x5-9 mm, usually ovate-rhomboid, flat, densely velutinous or glabrous; stems 20-30 cm; basal leaves obovate or obovate-oblanceolate subsp. cappadocica
1. Fruits 12-30x8-20 mm, elliptic, ovate, obovate or suborbicular, flat or undulate, puberulent or glabrous; stems 30-70 cm; basal leaves oblanceolate, often narrowly so
2. Wing of fruit strongly undulate when ripe
3. Fruit broadly elliptic to suborbicular, 13-17 x 9-17 mm subsp. steveniana
3. Fruit ovate-elliptic, 20-30 x 12-20 mm, often mottled with purplish brown subsp. macrocarpa
2. Wing of ripe fruit nearly flat
4. Fruits cuneate-oblong, 24-26 x8-9 mm, tapered below, wing as broad as loculus subsp. nurihakensis
4. Fruits suborbicular, obovate, ovate or elliptic, 12-30 x 9-20 mm, wing manifestly broader than loculus
5. Fruit suborbicular, 12-16 x 12-16 mm , obtuse or emarginate subsp. alyssifolia
5. Fruits obovate, elliptic or ovate-elliptic, 14-30 x9-20 mm
6. Fruits usually obovate or elliptic, 14-24 x 9-12 mm, acute at the base or at both ends; stem leaves usually obtusely auricled ssp subradiata
7. Fruits obovate, sometimes broadly elliptic , usually obtuse and less than 2 x longer than broad, puberulent or glabrous var. subradiata
7. Fruits elliptic, usually 2 x longer than broad, obtuse or emarginate, tomentose-puberulent or glabrous var. gudrunensis
6. Fruits ovate-elliptic, 20-30 x 12-20 mm, rounded at least at the base, often mottled with purplish brown; stem leaves subacutely auricled subsp. macrocarpa
1. Fruits 10-12x5-9 mm, usually ovate-rhomboid, flat, densely velutinous or glabrous; stems 20-30 cm; basal leaves obovate or obovate-oblanceolate subsp. cappadocica
1. Fruits 12-30x8-20 mm, elliptic, ovate, obovate or suborbicular, flat or undulate, puberulent or glabrous; stems 30-70 cm; basal leaves oblanceolate, often narrowly so
2. Wing of fruit strongly undulate when ripe
3. Fruit broadly elliptic to suborbicular, 13-17 x 9-17 mm subsp. steveniana
3. Fruit ovate-elliptic, 20-30 x 12-20 mm, often mottled with purplish brown subsp. macrocarpa
2. Wing of ripe fruit nearly flat
4. Fruits cuneate-oblong, 24-26 x8-9 mm, tapered below, wing as broad as loculus subsp. nurihakensis
4. Fruits suborbicular, obovate, ovate or elliptic, 12-30 x 9-20 mm, wing manifestly broader than loculus
5. Fruit suborbicular, 12-16 x 12-16 mm , obtuse or emarginate subsp. alyssifolia
5. Fruits obovate, elliptic or ovate-elliptic, 14-30 x9-20 mm
6. Fruits usually obovate or elliptic, 14-24 x 9-12 mm, acute at the base or at both ends; stem leaves usually obtusely auricled ssp subradiata
7. Fruits obovate, sometimes broadly elliptic , usually obtuse and less than 2 x longer than broad, puberulent or glabrous var. subradiata
7. Fruits elliptic, usually 2 x longer than broad, obtuse or emarginate, tomentose-puberulent or glabrous var. gudrunensis
6. Fruits ovate-elliptic, 20-30 x 12-20 mm, rounded at least at the base, often mottled with purplish brown; stem leaves subacutely auricled subsp. macrocarpa
